泰国晚始新世Anthracokeryx thailandicus Ducrocq, 1999(石炭兽科,小丘齿兽亚科)的分类修订
收稿日期: 2020-04-30
网络出版日期: 2020-10-19
基金资助
法国ANR-09-BLAN-0238项目;古灵长类和古人类及演化与古环境研究室和普瓦提埃大学资助
Taxonomic revision of Anthracokeryx thailandicus Ducrocq, 1999 (Anthracotheriidae, Microbunodontinae) from the Upper Eocene of Thailand
Received date: 2020-04-30
Online published: 2020-10-19
重新描述了泰国晚始新世石炭兽类(哺乳动物纲,鲸偶蹄类)的泰国先炭兽(Anthracokeryx thailandicus), 修订了该种在科内的系统发育位置。据观察,泰国标本具有一系列重要的牙齿差异,因而建立一新属颏炭兽(Geniokeryx gen. nov.), 代表了归入小丘齿兽亚科(Microbunodontinae)的第3个属。新属的主要特征是下颌联合短而深,未愈合;上、下前臼齿粗壮;上臼齿呈微弱新月型,具原附尖,无后小尖外棱。下颌联合的独特形态可能属于性双型特征,为雄性增大的上犬齿提供了侧面保护,就像某些古近纪祖猎虎类食肉动物,例如Eusmilus一样。简要评述了中国发现的先炭兽属的一些种,认为A. dawsoni可能是A. sinensis的同物异名。
DUCROCQ Stéphane . 泰国晚始新世Anthracokeryx thailandicus Ducrocq, 1999(石炭兽科,小丘齿兽亚科)的分类修订[J]. 古脊椎动物学报, 2020 , 58(4) : 293 -304 . DOI: 10.19615/j.cnki.1000-3118.200618
The anthracotheriid (Mammalia, Cetartiodactyla) species Anthracokeryx thailandicus from the Upper Eocene of Thailand is redescribed in details and a revision of its phylogenetic position within the family is proposed. A combination of important dental differences has been observed that led to attribute the Thai form to a distinct genus, Geniokeryx gen. nov., which represents the third genus included into the Microbunodontinae. The new genus is characterized mainly by its unfused short and deep mandibular symphysis, massive lower and upper premolars, weakly selenodont upper molars that exhibit a protostyle and lack an ectometacristule. The peculiar morphology of its symphysis might have been a sexually dimorphic feature that provided the role of a lateral protection for the enlarged upper canine in males as seen in some Paleogene nimravid carnivorans like Eusmilus. A short review of some Anthracokeryx species from China suggests that A. dawsoni might be synonymous to A. sinensis.
Key words: Thailand; Late Eocene; Anthracotheriidae; Microbunodontinae; Anthracokeryx; Geniokeryx
[1] | Averianov A, Obraztsova E, Danilov I et al., 2019. Anthracotheriid artiodactyl Anthracokeryx and an Upper Eocene age for the Youganwo Formation of southern China. Hist Biol, 31: 1115?1122 |
[2] | Boisserie J R, Lihoreau F, Orliac M et al., 2010. Morphology and phylogenetic relationships of the earliest known hippopotamids (Cetartiodactyla, Hippopotamidae, Kenyapotaminae). Zool J Linn Soc, 158:325-366 |
[3] | Ducrocq S, 1999. The Late Eocene Anthracotheriidae (Mammalia, Artiodactyla) from Thailand. Palaeontogr Abt A, 252:93-140 |
[4] | Ducrocq S, Benammi M, Chavasseau O et al., 2015. New anthracotheres (Cetartiodactyla, Mammalia) from the Paleogene of northeastern Vietnam: biochronological implications. J Vert Paleont, 35:e929139 |
[5] | Leidy J, 1869. The extinct mammalian fauna of Dakota and Nebraska, including an account of some allied forms from other localities, together with a synopsis of the mammalian remains of North America. J Acad Nat Sci Philadelphia, 7:1-472 |
[6] | Lihoreau F, Ducrocq S, 2007. Family Anthracotheriidae. In: Prothero D R, Foss S E eds. The Evolution of Artiodactyls. Baltimore: The Johns Hopkins University Press. 89?105 |
[7] | Lihoreau F, Blondel C, Barry J et al., 2004. A new species of the genus Microbunodon (Anthracotheriidae, Artiodactyla) from the Miocene of Pakistan: genus revision, phylogenetic relationships and palaeobiogeography. Zool Scr, 33: 97-115 |
[8] | Lihoreau F, Boisserie J R, Manthi F K et al., 2015. Hippos stem from the longest sequence of terrestrial cetartiodactyl evolution in Africa. Nat Commun, 6:6264 |
[9] | Montgelard C, Catzeflis F M, Douzery E, 1997. Phylogenetic relationships of artiodactyls and cetaceans as deduced from the comparison of cytochrome b and 12S rRNA mitochondrial sequences. Mol Biol Evol, 14(5):550-559 |
[10] | Pilgrim G E, Cotter G de P, 1916. Some newly discovered Eocene mammals from Burma. Rec Geol Surv India, 47:42-77 |
[11] | Qiu Z D, 1977. Notes on the new species of Anthracokeryx from Guangxi. Vert PalAsiat, 15(1):54-58 |
[12] | Soe A N, Chavasseau O, Chaimanee Y et al., 2017. New remains of Siamotherium pondaungensis (Cetartiodactyla, Hippopotamoidea) from the Eocene of Pondaung, Myanmar: paleoecologic and phylogenetic implications. J Vert Paleont, 37:e1312691 |
[13] | Swofford D L, 2002. PAUP*: Phylogenetic Analysis Using Parsimony (*And Other Methods), version 4. Sunderland, Massachusetts: Sinauer Associates |
[14] | Van Valkenburgh B, 2007. Déjà Vu: the evolution of feeding morphologies in the Carnivora. Integr Comp Biol, 47:147-163 |
[15] | Wang J W, 1985. A new species of Anthracokeryx from Shanxi. Vert PalAsiat, 23(1):58-59 |
[16] | Xu Y X, 1962. Some new anthracotheres from Shansi and Yunnan. Vert PalAsiat, 6(3):232-250 |
[17] | Zdansky O, 1930. Die altterti?ren S?ugetiere Chinas nebst stratigraphischen Bemerkungen. Palaeont Sin, 6(2):5-87 |
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