乳齿象类的命名与中文译法及分类学问题讨论
收稿日期: 2020-11-13
网络出版日期: 2021-07-28
基金资助
中国科学院战略性先导科技专项(XDB26000000);中国科学院战略性先导科技专项(XDA20070203);中国科学院战略性先导科技专项(XDB31030106);科技部基础性工作专项资助(2015FY310100)
On the scientific names of mastodont taxa: nomenclature, Chinese translation, and taxonomic problems
Received date: 2020-11-13
Online published: 2021-07-28
乳齿象类是长鼻类演化的重要阶段,该阶段奠定了长鼻类各冠群演化的基本格局。乳齿象类的研究有400多年的历史,分类和命名经过多次修订改动,乳齿象类的演化脉络完全体现在乳齿象命名历史之中。从词源学角度考证乳齿象类各类群的命名来源,整理了175条乳齿象类的中文译名(绝大多数为有效分类名), 包括12个属以上分类群,46个属,117个种,涵盖了乳齿象类几乎全部的种属。在此基础上梳理了乳齿象类的演化脉络,提出乳齿象类分类和命名中存在的一些问题。乳齿象类的头骨和下颌的演化在各支系中都是连续的,体现出了相近的平行演化趋势,而颊齿的形态特征虽然区别不明显,但在各支系中相对稳定。豕棱齿象科(Choerolophodontidae)是乳齿象类中最稳健的单系群,其中厚棱象(Synconolophus )可能是有效属名;玛姆象科中,中新乳齿象(Miomastodon )和上新乳齿象(Pliomastodon )可能都是有效的,但不一定是美洲乳齿象(Mammut americanum )的直接祖先;铲齿象科(Amebelodontidae)中达氏铲齿象(Platybelodon danovi )与格氏铲齿象(P. grangeri )、赵氏隐门齿象(Aphanobelodon zhaoi )的系统发育关系存在疑问,取决于P. danovi 的下门齿断面究竟是否为齿柱状结构,而美洲的布氏柱门齿象(Konobelodon britti )可能是卢氏匙门齿象(Torynobelodon loomisi )的同物异名,亚洲归入Konobelodon 的种不一定是铲齿象类,可能应归为副四棱齿象(Paratetralophodon ); 嵌齿象科(Gomphotheriidae)中锯齿象属(Serridentinus )可能有效,它代表了嵌齿象类中一个偏轭型化的类群,向居维叶象亚科方向演化;居维叶象亚科(Cuvieroniinae)可能仅包括居维叶象(Cuvieronius )和喙嘴象(Rhynchotherium )属,而美洲其他的短颌嵌齿象类中,脊乳齿象(Stegomastodon )有可能从铲齿象科中的一支演化而来,南方乳齿象(Notiomastodon )则可能与中华乳齿象(Sinomastodon )相关;Sinomastodon 可能起源于中国南方的竹棚上新乳齿象(Pliomastodon (?) zhupengensis ), 原来的属型种中间中华乳齿象(Sinomastodon intermedius )具有早出原同名,建议以它的早出异名仙台中华乳齿象(Sinomastodon sendaicus )取代S. intermedius 。
王世骐, 李春晓, 张晓晓 . 乳齿象类的命名与中文译法及分类学问题讨论[J]. 古脊椎动物学报, 2021 , 59(4) : 295 -332 . DOI: 10.19615/j.cnki.2096-9899.210728
The mastodont-grade proboscideans represent an important stage in the evolution of the group, establishing the basic pattern of the evolution of the crown groups of proboscideans. The research on mastodons has a history of more than 400 years. The classification and nomenclature have been revised and changed many times, and the problems in their evolution were fully reflected in the history of mastodon nomenclature. In this paper, we undertook a bibliographical research into the nomenclature and etymology of various mastodont groups, reviewing 175 translated Chinese names of mastodont-grade proboscideans, including 12 taxon names higher than the genus level, 46 genera, and 117 species, covering almost all the species of the mastodont radiation. On this basis, we review the principal phylogenetic hypotheses of mastodont interrelationships, and highlight problems in the classification and nomenclature of mastodonts. The evolution of the skull and mandible of mastodons is continuous in all clades, reflecting the same parallel evolution trend; while, although the morphological characteristics of cheek teeth across all lineages are not obvious, they are relatively stable in each lineage. Choerolophodontidae is the most robust monophyletic group within the mastodonts, of which Synconolophus may be a distinct, valid genus. Miomastodon and Pliomastodon of Mammutidae may both be valid, but they are not necessarily the direct ancestor of Mammut americanum . The phylogenetic relationship between Platybelodon danovi , P. grangeri and Aphanobelodon zhaoi within the Amebelodontidae is questionable, depending on whether the lower incisor section of P. danovi is the dentine rod structure or not, while Konobelodon britti in America may be a synonym of Torynobelodon loomisi . The species assigned to Konobelodon in Asia is possibly not amebelodontids, but probably attributable to Paratetralophodon , instead; Serridentinus of Gomphotheriidae may be a valid taxon, representing a trend towards somewhat zygodonty in Gomphotheriidae that terminated with the Cuvieroniinae. The Cuvieroniinae may only include Cuvieronius and Rhynchotherium , while other brevirostrine gomphotheres in America, such as Stegomastodon may have been evolved from a lineage of amebelodonts. Notiomastodon may be related to Sinomastodon , which itself may have originated from Pliomastodon (?) zhupengensis in southern China. The name Mastodon intermedius Teilhard de Chardin & Trassaert, 1937 (now Sinomastoodon intermedius ) has the senior primary homonym Mastodon intermedius Eichwald, 1831. We suggest that Sinomastodon intermedius should be replaced with its senior synonym-Sinomastodon sendaicus (
Key words： mastodons; proboscidean; etymology; nomemclature; Chinese translation; taxonomy
| [1] | Ameghino F, 1888. Rápidas diagnosis de algunos Mamíferos Fósiles Nuevos de la República Argentina. Buenos Aires: P E Coni. 1-17 |
| [2] | Andrews C W, 1901. Preliminary note on some recently discovered extinct vertebrates from Egypt. Part I. Geol Mag New Ser, 8:400-409 |
| [3] | Andrews C W, Beadnell H J L, 1902. A Preliminary Note on Some New Mammals from the Upper Eocene of Egypt. Cairo: Survey Department, Public Works Ministry. 1-9 |
| [4] | Arambourg C, 1945. Un Mastodonte nouveau du Pliocène inférieur d’Égypte. Bull Soc Géol Fr, 5:479-495 |
| [5] | Aymard A, 1855. Anancus Aymard, Anancus macroplus Aymard. In: Dorlhac M J ed. Notice geologique sur le cratere de Coupet et sur son gisement de gemmes et d’ossements fossils. Ann Soc Agr, Sci, Arts Comm Puy, 19:497-517 |
| [6] | Barbour E H, 1914. A new longirostral mastodon from Cherry County, Nebraska. Nebr Geol Surv, 4:213-222 |
| [7] | Barbour E H, 1916. A new longirostral mastodon from Nebraska, Tetrabelodon osborni , sp. nov. Am J Sci, 41:522-529 |
| [8] | Barbour E H, 1927. Preliminary notice of a new proboscidean Amebelodon fricki , gen. et sp. nov. Bull Nebr State Mus, 1:131-134 |
| [9] | Barbour E H, 1929. Torynobelodon loomisi , gen. et sp. nov. Bull Nebr State Mus, 16:147-153 |
| [10] | Barbour E H, 1931. A new amebelodont, Torynobelodon barnumbrowni , sp. nov., a preliminary report. Bull Nebr State Mus, 22:191-198 |
| [11] | Barbour E H, Sternberg G F, 1935. Gnathabelodon thorpei , gen. et sp. nov., a new mud-grubbing mastodon. Bull Nebr State Mus, 42:395-403 |
| [12] | Bergounioux F M, Crouzel F, 1960. Tetralophodon curvirostris n. sp. (Mammalia, Proboscidea) aus dem Unterpliozän (Pontien) von Esselborn (Rheinhessen). Jahresb Mitt Oberrhein Geol Verein, 42:109-121 |
| [13] | Blumenbach J F, 1799. Handbuch der Naturgeschichte, 6th ed. Göttingen: Bei Heinrich Dieterich. 1-814 |
| [14] | Borissiak A A, 1928. On a new mastodon from the Chokrak beds (Middle Miocene) of the Kuban region, Platybelodon danovi n. gen. n. sp. Ann Paleontol Soc Russ, 7:105-120 |
| [15] | Borissiak A A, 1936. Mastodon atavus n. sp., der primitivste vertreter der gruppe M. angustidens . Trav Inst Paleozool Acad Sci URSS, 5:171-234 |
| [16] | Borissiak A A, Belyaeva E, 1928. Trilophodon (Serridentinus ?) inopinatus n. sp. from the Jilančik Beds of the Turgai Region. Bull Acad Sci URSS, Classe Sci Phys-Math, 1928:241-252 |
| [17] | Brown R W, 1954. Composition of Scientific Words. Washington, DC: Smithsonian Institution Press. 1-882 |
| [18] | Buckley M, Recabarren O P, Lawless C et al., 2019. A molecular phylogeny of the extinct South American gomphothere through collagen sequence analysis. Quat Sci Rev, 224:e105882 |
| [19] | Burmeister H, 1837. Handbuch der Naturgeschichte. Zum Gebrauch bei Vorlesungen. Berlin: Zweite Abteilung, Zoologie, T. C. F. Enslin. 1-795 |
| [20] | Cabrera Á, 1929. Una revisión de los mastodontes Argentions. Rev Mus Plata, 32:61-144 |
| [21] | Cautley P T, 1836. Note on the teeth of the Mastodonte à dents etroites of Siwálik Hills. J Asiat Soc Bengal, 5:294-296 |
| [22] | Chang X Z, Zhai R J, 1978. Miocene mastodonts of Lantian and Lintung, Shensi. Prof Pap Stratigr Palaeontol, 7:136-142 |
| [23] | Chen G F, 1978. Mastodont remains form the Miocene of Zhongning-Tongxin region in Ningxia. Vert PalAsiat, 16:103-110 |
| [24] | Chow M C, 1959. New species of fossil Proboscidea from South China. Acta Palaeontol Sin, 7:251-258 |
| [25] | Chow M C, Chang Y P, 1961. New mastodonts from North China. Vert PalAsiat, (3):245-255 |
| [26] | Chow M C, Chang Y P, 1974. Chinese Fossil Elephantoids. Beijing: Science Press. 1-74 |
| [27] | Clift W, 1828. On the fossil remains of two new species of mastodon, and of other vertebrated animails, found on the left bank of the Irawadi. Trans Geol Soc London, 2:369-375 |
| [28] | Colbert E H, 1969. Evolution of the Vertebrates. A History of the Backboned Animals through Time. 2nd ed. Translated by Zhou M Z, Liu H Y, Zhou B X, 1976. Beijing: Geological Publishing House. 1-518 |
| [29] | Cope E D, 1873. Synopsis of New Vertebrata from the Tertiary of Colorado, Obtained During the Summer of 1873. Washington: Government Printing Office. 1-19 |
| [30] | Cope E D, 1875. On a new mastodon and rodent. Proc Acad Nat Sci Phila, 26:221-223 |
| [31] | Cope E D, 1878. Descriptions of new vertebrate from the Upper Tertiary Formations of the West. Proc Am Phil Soc, 17:219-231 |
| [32] | Cope E D, 1884. The extinct mammalian of the valley of Mexico. Proc Am Phil Soc, 22:1-21 |
| [33] | Coppens Y, 1972. Un nouveau proboscidien de Pliocene du Tchad, Stegodibelodon schneideri nov. gen. nov. sp., et le phylum des Stegotetrabelodontinae. C R Séances Acad Sci, 274:2962-2965 |
| [34] | Croizet J B, Jobert A C G, 1828. Recherches sur les ossemens fossiles du Département du Puy-De-Dôme. Paris: Principaux Libraires. 1-224 |
| [35] | Cuvier G, 1806. Sur différentes dents du genre des Mastodontes, mais d’espèces moindres que celles de l’Ohio, trouvées en plusieurs lieux des deux continents. Ann Mus Hist Nat, 8:401-424 |
| [36] | Cuvier G, 1817. Le règne animal. Vol. 1. Paris: Dé terville. 1-642 |
| [37] | Depéret C, 1897. Découverte du Mastodon angustidens dans l’étage cartennein de Kabylie. Bull Soc Géol Fr, 3:518-521 |
| [38] | Desmarest A G, 1822. Mammalogie ou Description des Espèces de Mammifère. Seconde Partie. Paris: Veuve Agasse. 1-555 |
| [39] | Dooley Jr A C, Scott E, Green J et al., 2019. Mammut pacificus sp. nov., a newly recognized species of mastodon from the Pleistocene of western North America. Peer J, 7:1-58 |
| [40] | Eichwald E, 1831. Zoologia specialis III. Vilnius: Josophi Zawadski. 1-404 |
| [41] | Eichwald E, 1835. De pecorum et pachydermorum reliquiis fossilibus in Lithuania, Volhynia et Podolia repertis. Nova Acta Acad C Leopold Carol, 17:677-780 |
| [42] | Falconer H, 1857. On the species of mastodon and elephant occurring in the fossil state in Great Britain. Part I. Mastodon . Quart J Geol Soc London, 13:307-360 |
| [43] | Falconer H, 1868. Paleontological Memoirs and Notes of the Late Hugh Falconer with a Biographical Sketch of the Author. London: Hardwicke. 1-590 |
| [44] | Falconer H, Cautley P T, 1846. Fauna Antiqua Sivalensis, Being the Fossil Zoology of the Siwalik Hills, in the North of India. London: Smith, Elder. 1-64 |
| [45] | Fischer de Waldheim G, 1814. Zoognosia. Tabulis synopticis illustrata, in uswn Praelectionwn Academiae Imperialis Medicochirurgicae. Mosque: Mosquensis Edita, Nicolai Sergeidis Vsevolozsky. 1-694 |
| [46] | Fourtau R, 1918. Contribution a l’etude des vertebres miocenes de l’Egypte. Ministry of Finance. Cairo: Survey Department, Government Press. 1-99 |
| [47] | Frick C, 1926. Tooth sequence in certain trilophodont tetrabelodont mastodons and Trilophodon (Serridentinus ) pojoaquensis , new species. Bull Am Mus Nat Hist, 56:122-178 |
| [48] | Frick C, 1933. New remains of trilophodont-tetrabelodont mastodons. Bull Am Mus Nat Hist, 56:505-652 |
| [49] | Gaudry A, Lartet E, 1856. Résultats des recherches paléontologiques entreprises dans l’Attique sous les auspices de l’Académie. C R Acad Sci, 43:271-274 |
| [50] | Gaziry A W, 1976. Jungtertiäre Mastodonten aus Anatolien (Türkei). Geol Jahrb, 22:3-143 |
| [51] | Gaziry A W, 1997. Die Mastodonten (Proboscidea, Mammalia) aus Dorn-Dürkheim 1 (Rheinhessen). In: Franzen J L ed. Die Säuteqiere dus dem Turolium von Dorn-Dürkheim 1 (Rheinhessen, Deutschland). Cour Forschungsinst Senckenberg, 197:73-115 |
| [52] | Gheerbrant E, Tassy P, 2009. L’origine et l’évolution des éléphants. C R Palevol, 8:281-294 |
| [53] | Gray J E, 1821. On the natural arrangement of vertebrose animals. London Med Repos, 15:296-310 |
| [54] | Guan J, 1988. The Miocene strata and mammals from Tongxin, Ningxia and Guanghe, Gansu. Mem Beijing Nat Hist Mus, 42:1-21 |
| [55] | Hay O P, 1922. Further observations on some extinct elephants. Proc Biol Soc Wash, 35:97-101 |
| [56] | Hays I, 1834. Descriptions of the specimens of inferior maxillary bones of mastodonts in the Cabinet of the American Philosophical Society, with remarks on the Genus Tetracaulodon (Godman) etc. Trans Am Philos Soc, 4:317-339 |
| [57] | Hoffstetter R, 1950. Observaciones sobre los mastodontes de Sud América y especialmente del Ecuador: Haplomastodon , subgn. nov. de Stegomastodon . Publ Esc Politec Nac, 1:1-51 |
| [58] | Hopwood A T, 1935. Fossil Proboscidea from China. Palaeontol Sin, Sér C, 9:1-108 |
| [59] | Ji X P, Zhang X Y, 1997. Proboscidea. In: He Z Q ed. Yuanmou Hominoid Fauna. Kunming: Yunnan Science and Technology Press. 89-93 |
| [60] | Kaup J J, 1835. Description d’Ossements fossiles de Mammifères, Cahier IV. In: Heyer J G ed. Inconnus jusqu’a présent, qui se trouvent au Musée grand-ducal de Darmstadt. Darmstadt: Hardpress Publishing. 65-89 |
| [61] | Kerr R, 1792. The Animal Kingdom or Zoological System, of the Celebrated Sir Charles Linnaeus; Class I. Mammalia. Edingburgh: Printed for A. Strahan, and T. Cadell, London, and W. Creech. 1-680 |
| [62] | Klähn H, 1931. Rheinhessisches Pliozän, im Rahmen des mitteleuropäischen Pliozäns. Geol Palaeontol, 18:279-339 |
| [63] | Koenigswald G H R v, 1959. A mastodon and other fossil mammals from Thailand. Rep Invest R Depart Min, 2:25-28 |
| [64] | Konidaris G E, Roussiakis S J, Theodorou G E et al., 2014. The Eurasian occurrence of the shovel-tusker Konobelodon (Mammalia, Proboscidea) as illuminated by its presence in the Late Miocene of Pikermi (Greece). J Vert Paleont, 34(6):1437-1453 |
| [65] | Lambert W D, 1990. Rediagnosis of the genus Amebelodon (Mammalia, Proboscidea, Gomphotheriidae), with a new subgenus and species, Amebelodon (Konobelodon ) britti . J Paleontol, 64:1032-1040 |
| [66] | Lambert W D, 2007. New tetralophodont gomphothere material from Nebraska and its implications for the status of North American Tetralophodon . J Vert Paleont, 27:676-682 |
| [67] | Lambert W D, 2016. Eurybelodon shoshanii , an unusual new shovel-tusked gomphothere (Mammalia, Proboscidea) from the Late Miocene of Oregon. J Vert Paleont, 36:1-7 |
| [68] | Leidy J, 1858. On new species of mastodon and elephant from Nebraska, Mastodon mirificus , Elephas imperator . Proc Acad Nat Sci Philadelphia, 10:1-10 |
| [69] | Leidy J, 1886. Mastodon and Llama from Florida. Proc Acad Nat Sci Philadelphia, 38:11-12 |
| [70] | Li C X, Wang S Q, Mothé D et al., 2019. New fossils of Early and Middle Miocene Choerolophodon from northern China reveal a Holarctic distribution of Choerolophodontidae. J Vert Paleont, doi: 10.1080/02724634.2019.1618864 |
| [71] | Li C X, Ji X P, Zhang S T et al., 2021. The new fossil record of Stegolophodon latidens from the Xiaolongtan locality, Yunnan, China, and the discussion on the age of the Lufengpithecus keiyuanensis . Chinese Sci Bull, 66:1469-1481, doi: 10.1360/TB-2020-0769 |
| [72] | Lofgren D L, Anand R S, 2011. Partial skull of Zygolophodon (Mammalia, Proboscidea) from the Barstow Formation of California. J Vert Paleont, 31:1392-1396 |
| [73] | Lucas S G, Morgan G S, 2008a. Taxonomy of Rhynchotherium (Mammalia, Proboscidea) from the Miocene-Pliocene of North America. New Mex Mus Nat Hist Sci, 44:70-88 |
| [74] | Lucas S G, Morgan G S, 2008b. The proboscidean Amebelodon from east-central New Mexico and its biochronological significance. New Mex Mus Nat Hist Sci, 44:89-92 |
| [75] | Lydekker R, 1884. Additional Siwalik Perissodactyla and Proboscidea. Mem Geol Surv India, Palaeont Indica, 10(3):1-34 |
| [76] | Lydekker R, 1886. Addenda to synopsis of Siwalik & Narbada Mammalia. Mem Geol Surv India, Palaeont Indica, 10(4):1-22 |
| [77] | Maarel F H van der, 1932. Contribution to the knowledge of the fossil mammalian fauna of Java. Wet Med, Dienst Mijnbouw Nederland-Indiё, 15:1-208 |
| [78] | MacInnes D G, 1942. Miocene and post-Miocene Proboscidea from east Africa. Trans Zool Soc London, 25:33-106 |
| [79] | Maglio V J, 1970. Four new species of Elephantidae from the Plio-Pleistocene of northwestern Kenya. Breviora, 341:1-43 |
| [80] | Maglio V J, 1974. A new proboscidean from the Late Miocene of Kenya. Palaeontology, 17:699-705 |
| [81] | Markov G N, 2008. The Turolian proboscideans (Mammalia) of Europe: preliminary observations. Hist Nat Bulg, 19:153-178 |
| [82] | Matsumoto H, 1924. Prelimenary notes on two new species of fossil mastodon from Japan. J Geol Soc Tokyo, 31:395-414 |
| [83] | Matthew W D, 1918. Contributions to the Snake Creek fauna: with notes upon the Pleistocene of western Nebraska-American Expedition of 1916. Bull Am Mus Nat Hist, 38:183-229 |
| [84] | Mayet L, 1908. Étude des mammifères Miocènes des sables de l’Orléanais et des faluns de la Touraine. Ann Univ Lyon, Nouv Sér I, Sci, Méd, 24:1-321 |
| [85] | Mothé D, Avilla L, 2015. Mythbusting evolutionary issues on South American Gomphotheriidae (Mammalia: Proboscidea). Quat Sci Rev, 110:23-35 |
| [86] | Mothé D, Avilla L S, Zhao D et al., 2016a. A new Mammutidae (Proboscidea, Mammalia) from the Late Miocene of Gansu Province, China. An Acad Bras Ciênc, 88:65-74 |
| [87] | Mothé D, Ferretti M P, Avilla L S, 2016b. The dance of tusks: rediscovery of lower incisors in the Pan-American proboscidean Cuvieronius hyodon revises incisor evolution in Elephantimorpha. PLoS One, 11:e0147009 |
| [88] | Mothé D, Ferretti M P, Avilla L S, 2019. Running over the same old ground: Stegomastodon never roamed South America. J Mamm Evol, 26:165-177 |
| [89] | Mucha B B, 1980. A new species of ridge-toothed mastodon from the Pliocene of south-western USSR. In: Negadaev-Nikonov K N ed. Quaternary and Neogene Faunas and Flora of the Moldavian SSR. Shtiintza: Kishinev. 19-26 |
| [90] | Osborn H F, 1918. A long-jawed mastodon skeleton from South Dakota and phylogeny of the Proboscidea. Bull Geol Soc Am, 29:133-137 |
| [91] | Osborn H F, 1921. First appearance of the true Mastodon in America. Am Mus Novit, 10:1-6 |
| [92] | Osborn H F, 1922. Dibelodon edensis (Frick) of southern California, Miomastodon of the Middle Miocene, new genus. Am Mus Novit, 49:1-4 |
| [93] | Osborn H F, 1923. New subfamily, generic, and specific stage in the evolution of the Proboscidea. Am Mus Novit, 99:1-4 |
| [94] | Osborn H F, 1924a. Additional generic and specific stages in the evolution of the Proboscidea. Am Mus Novit, 154:1-5 |
| [95] | Osborn H F, 1924b. Serridentinus and Baluchitherium , Loh Fomation, Mongolia. Am Mus Novit, 148:1-5 |
| [96] | Osborn H F, 1926. Additional new genera and species of the mastodontoid Proboscidea. Am Mus Novit, 238:1-16 |
| [97] | Osborn H F, 1929a. New Eurasiatic and American proboscideans. Am Mus Novit, 393:1-23 |
| [98] | Osborn H F, 1929b. The revival of Central Asiatic life. Nat Hist, 29:3-16 |
| [99] | Osborn H F, 1932. Trilophodon cooperi n. sp. of Dera Bugti, Baluchistan. Am Mus Novit, 393:1-6 |
| [100] | Osborn, H F, 1936. Proboscidea: a Monograph of the Discovery, Evolution, Migration and Extinction of the Mastodonts and Elephants of the World (Vol. I). New York: The American Museum Press. 1-802 |
| [101] | Osborn H F, Granger W, 1932. Platybelodon grangeri , three growth stages, and a new serridentine from Mongolia. Am Mus Novit, 537:1-13 |
| [102] | Ozansoy F, 1965. Étude des gisements continentaux et des mammifères du Cénozoïque de Turquie. Mém Soc Géol Fr, 102:1-92 |
| [103] | Petrocchi C, 1943. Il giacimento fossilifero di Sahabi. Coll Sci e Doc. Verbania: Ministero dell’Africa Italiana. 1-162 |
| [104] | Pickford M, 1987. Révision des suiformes (Artiodactyla, Mammalia) de Bugti (Pakistan). Ann Paléont, 73:289-350 |
| [105] | Pickford M, 2001. Afrochoerodon nov. gen. kisumuensis (MacInnes) (Proboscidea, Mammalia) from Cheparawa, Middle Miocene, Kenya. Ann Paléont, 87:99-117 |
| [106] | Pickford M, 2003. New Proboscidea from the Miocene strata in the lower Orange River Vally, Namibia. Mem Geol Surv Namibia, 19:207-256 |
| [107] | Pickford M, Tassy P, 1980. A new species of Zygolophodon (Mammalia, Proboscidea) from the Miocene hominoid localities of Meswa Bridge and Moroto (East Africa). Neues Jahrb Geol Paläont Monatsh, 4:235-251 |
| [108] | Pilgrim G E, 1908. The Tertiary and post-Tertiary freshwater deposits of Baluchistan and Sind with notices of new vertebrates. Rec Geol Surv India, 37:139-166 |
| [109] | Pilgrim G E, 1912. The vertebrate fauna of the Gaj Series in the Bugti Hills and the Punjab. Mem Geol Surv India,Palaeont Indica, 4:1-83 |
| [110] | Pilgrim G E, 1913. The correlation of the Siwaliks with mammal horizons of Europe. Rec Geol Surv India, 43:264-326 |
| [111] | Pohlig H, 1912. Sur une vieille mandibule de “Tetracaulodon ohiotocum ” Blum., avec défense in situ. Bull Soc Bélg Geol, 26:187-193 |
| [112] | Rasmussen D T, Gutierrez M, 2009. A mammalian fauna from the Late Oligocene of northwestern Kenya. Palaeontogr Abt A: Paläozool, Stratigr, 288:1-52 |
| [113] | Saegusa H, Thasod Y, Ratanasthien B, 2005. Notes on Asian stegodontids. Quat Int, 126- 128:31-48 |
| [114] | Sanders W J, Miller E R, 2002. New proboscideans from the Early Miocene of Wadi Moghara, Egypt. J Vert Paleont, 22:388-404 |
| [115] | Sanders W J, Gheerbrant E, Harris J M, et al., 2010. Proboscidea. In: Werdelin L, Sanders W J eds. Cenozoic Mammals of Africa. Berkeley: University of California Press. 161-251 |
| [116] | Schinz H R, 1824. Naturgeschichte und Abbildungen der Säugethiere. Zurich: Brodtmannx, Lithographischer Kunstanstalt. 1-417 |
| [117] | Schlesinger G, 1917. Die Mastodonten des K. K. Naturhistorischen Hofmuseums. Denkschr K K Naturhist Hofmus 1,Geol Paläont Reihe, 1:1-231 |
| [118] | Schlesinger G, 1919. Die stratigraphische Bedeutung der europäischen Mastodonten. Mitt Geol Gesell Wien, 11:129-166 |
| [119] | Sellards E H, 1940. New Pliocene mastodon. Bull Geol Soc Am, 51:1659-1664 |
| [120] | Shoshani J, Tassy P, 1996. The Proboscidea: Evolution and Palaeoecology of Elephants and Their Relatives. Oxford: Oxford University Press. 1-472 |
| [121] | Shoshani J, Tassy P, 2005. Advances in proboscidean taxonomy & classification, anatomy & physiology, and ecology & behavior. Quat Int, 126- 128:5-20 |
| [122] | Shoshani J, Golenberg G M, Yang H, 1998. Elephantidae phylogeny: morphological versus molecular results. Acta Theriol, 5:89-122 |
| [123] | Shoshani J, Walter R C, Abraha M et al., 2006. A proboscidean from the Late Oligocene of Eritrea, a “missing link” between early Elephantiformes and Elephantimorpha, and biogeographic implications. Proc Nat Acad Sci, 103:17296-17301 |
| [124] | Simpson G G, 1945. The principles of classification and a classification of mammals. Bull Am Mus Nat Hist, 85:1-350 |
| [125] | Tang Y J, Zong G F, Lei Y L et al., 1987. Mammalian remains from the Pliocene of the Hanshui River Basin, Shaanxi. Vert PalAsiat, 25:222-235 |
| [126] | Tassy P, 1977. Découverte de Zygolophodon turicensis (Schinz) (Proboscidea, Mammalia) au lieu-dit Malartic a Simorre, Gers (Vindobonien moyen): implications paléoécologiques et biostratigraphiques. Geobios, 10:655-669 |
| [127] | Tassy P, 1983a. Les Elephantoidea Miocènes du Plateau du Potwar, Groups de Siwalik, Pakistan. IIe Partie: choerolophodontes et gomphothères. Ann Paléont, 69:235-297 |
| [128] | Tassy P, 1983b. Les Elephantoidea Miocènes du Plateau du Potwar, Groups de Siwalik, Pakistan. IIIe Partie: stégodontidés éléphantoïdes indéterminés restes postcrâniens. Conclusions. Ann Paléont, 69:317-354 |
| [129] | Tassy P, 1984. Le mastodonte à dents étroites, le grade trilophodonte et la radiation initiale des Amebelodontinae. In: Buffetaut E, Mazin J M, Salmon E eds. Actes du Symposium Paléontologique Georges Cuvier. Montbéliard: Impressions le Serpentaire. 459-473 |
| [130] | Tassy P, 1985. La place des mastodontes Miocènes de l’ancien monde dans la phylogénie des Proboscidea (Mammalia): hypothèses et conjectures.(Vol. I-III).Thèse Doctorat ès Sciences. Paris: Université Pierre et Marie Curie. 1-861 |
| [131] | Tassy P, 1988. The classification of Proboscidea: how many cladistic classifications? Cladistics, 4:43-57 |
| [132] | Tassy P, 2016. Proboscidea. In: Sen S ed. Late Miocene Mammal Locality of Küçükçekmece, European Turkey. Geodiversitas, 38:261-271 |
| [133] | Tassy P, Pickford M, 1983. Un nouveau mastodonte zygolophodonte (Proboscidea, Mammalia) dans le Miocène inférieur d’Afrique orientale: systématique et paléoenvironnement. Géobios, 16:53-77 |
| [134] | Tassy P, Anupandhanant P, Ginsburg L et al., 1992. A new Stegolophodon (Proboscidea, Mammalia) from the Early Miocene of northern Thailand. Géobios, 25:511-523 |
| [135] | Tassy P, Goussard F, Sanz M G, 2013. The status of Mastodon angustidens pygmaeus Depéret, 1897 (Proboscidea, Mammalia): the contribution of X-ray tomography. Géobios, 46:329-334 |
| [136] | Teilhard de Chardin P, Trassaert M, 1937. The proboscidians of south-eastern Shansi. Palaeont Sin Ser C, 13:1-58 |
| [137] | Tobien H, 1972. Status of the genus Serridentinus Osborn 1923 (Proboscidea, Mammalia) and related forms. Mainzer Geowiss Mitt, 1:143-191 |
| [138] | Tobien H, 1973. The structure of the mastodont molar (Proboscidea, Mammalia). Part 1: the bunodont patterns. Mainzer Geowiss Mitt, 2:115-147 |
| [139] | Tobien H, 1975. The structure of the mastodont molar (Proboscidea, Mammalia). Part 2: the zygodont and the zygobunodont patterns. Mainzer Geowiss Mitt, 4:195-233 |
| [140] | Tobien H, 1978. The structure of the mastodont molar (Proboscidea, Mammalia). Part 3: the Oligocene mastodont genera Palaeomastodon , Phiomia and the Eo/Oligocene paenungulate Moeritherium . Mainzer Geowiss Mitt, 6:177-208 |
| [141] | Tobien H, 1980. A note on the skull and mandible of a new choerolophodont mastodont (Proboscidea, Mammalia) from the Middle Miocene of Chios (Aegean Sea, Greece). In: Jacobs L ed. Aspects of Vertebrate History: Essays in Honor of Edwin Harris Colbert. Flagstaff: Museum of Northern Arizona Press. 299-307 |
| [142] | Tobien H, Chen G F, Li Y Q, 1986. Mastodonts (Proboscidea, Mammalia) from the Late Neogene and Early Pleistocene of the People’s Republic of China, part I: historical account: the genera Gomphotherium , Choerolophodon , Synconolophus , Amebelodon , Platybelodon , Sinomastodon . Mainzer Geowiss Mitt, 15:119-181 |
| [143] | Tobien H, Chen G F, Li Y Q, 1988. Mastodonts (Proboscidea, Mammalia) from the Late Neogene and Early Pleistocene of the People’s Republic of China, part II: the genera Tetralophodon , Anancus , Stegotetrabelodon , Zygolophodon , Mammut , Stegolophodon . Mainzer Geowiss Mitt, 17:95-220 |
| [144] | Tomida Y, Nakaya H, Saegusa H , et al., 2013. Miocene land mammals and stratigraphy of Japan. In: Wang X M, Flynn L J, Fortelius M eds. Neogene Terrestrial Mammalian Biostratigraphy and Chronology of Asia. New York: Columbia University Press. 314-333 |
| [145] | Translation Information Group of Xinhua News Agency, 1989. English Name Translation Manual. Beijing: The Commercial Press. 1-488 |
| [146] | Vacek V M, 1877. Über österreichische Mastodonten und ihre Beziehungen zu den Maston-Arten Europas. Abh Kaiserlich-Königlichen Geol Reichsanstalt, 7:1-45 |
| [147] | Wagner A, 1857. Neue Beitriige sur Kenntniss der fossilen Säugthier-Ueberreste von Pikermi. Abh Bayer Akad Wiss, 8:111-158 |
| [148] | Wang B Y, Qiu Z X, 2002. A new species of Platybelodon (Gomphotheriidae, Proboscidea, Mammalia) from Early Miocene of the Danghe area, Gansu, China. Vert PalAsiat, 40:291-299 |
| [149] | Wang S Q, Deng T, 2011. The first Choerolophodon (Proboscidea, Gomphotheriidae) skull from China. Sci China Earth Sci, 54:1326-1337 |
| [150] | Wang S Q, Deng T, Tang T et al., 2015a. Evolution of Protanancus (Proboscidea, Mammalia) in East Asia. J Vert Paleont, 35:1-13 |
| [151] | Wang S Q, Duangkrayom J, Yang X W, 2015b. Occurrence of the Gomphotherium angustidens group in China, based on a revision of Gomphotherium connexum (Hopwood, 1935) and Gomphotherium shensiensis Chang and Zhai, 1978: continental correlation of Gomphotherium species across the Palearctic. Paläont Z, 89:1073-1086 |
| [152] | Wang S Q, Ji X P, Jablonski N G et al., 2016a. The oldest cranium of Sinomastodon (Proboscidea, Gomphotheriidae), discovered in the uppermost Miocene of southwestern China: implications for the origin and migration of this taxon. J Mamm Evol, 23:155-173 |
| [153] | Wang S Q, Shi Q Q, He W et al., 2016b. A new species of the tetralophodont amebelodontine Konobelodon Lambert, 1990 (Proboscidea, Mammalia) from the Late Miocene of China. Geodiversitas, 38:65-97 |
| [154] | Wang S Q, Li Y, Duangkrayom J et al., 2017a. Early Mammut from the Upper Miocene of northern China, and its implications for the evolution and differentiation of Mammutidae. Vert PalAsiat, 55:233-256 |
| [155] | Wang S Q, Deng T, Ye J et al., 2017b. Morphological and ecological diversity of Amebelodontidae (Proboscidea, Mammalia) revealed by a Miocene fossil accumulation of an upper-tuskless proboscidean. J Syst Palaeont, 15:601-615 |
| [156] | Wang S Q, Li Y, Duangkrayom J et al., 2017c. A new species of Gomphotherium (Proboscidea, Mammalia) from China and the evolution of Gomphotherium in Eurasia. J Vert Paleont, 37:1-15 |
| [157] | Wang S Q, Saegusa H, Duangkrayom J et al., 2017d. A new species of Tetralophodon from the Linxia Basin and the biostratigraphic significance of tetralophodont gomphotheres from the Upper Miocene of northern China. Palaeoworld, 26:703-717 |
| [158] | Wang S Q, Ji X P, Deng T et al., 2019. Yunnan, a refuge for trilophodont proboscideans during the Late Miocene aridification of East Asia. Palaeogeogr Palaeoclimatol Palaeoecol, 515:162-171, doi: 10.1016/j.palaeo.2017.07.034 |
| [159] | Wang S Q, Zhang X X, Li C X, 2020. Reappraisal of Serridentinus gobiensis Osborn & Granger and Miomastodon tongxinensis Chen: the validity of Miomastodon . Vert PalAsiat, 58:134-158 |
| [160] | Wang Y, Jin C Z, Deng C L et al., 2012. The first Sinomastodon (Gomphotheriidae, Proboscidea) skull from the Quaternary in China. Chinese Sci Bull, 57:4726-4734 |
| [161] | Welcomme J L, 1994. Le plus ancien crâne de proboscidien d’Europe, Gomphotherium hannibali nov. sp. (Proboscidea, Mammalia), du Miocène inférieur du Languedoc (France). C R Acad Sci Sér II, 319:135-140 |
| [162] | Xiao D R, Zhou D G, Guo G R, 1984. The Manual of Boundaries and Toponym. Beijing: Encyclopedia of China Publishing House. 1-2806 |
| [163] | Xin H, 1970. French Name Translation Manual. Beijing: China Industry Press. 1-203 |
| [164] | Yabe H, 1950. Three alleged occurrences of Stegolophodon latidens (Clift) in Japan. Proc Jpn Acad, 26:61-65 |
| [165] | Yang X W, Li Y, Wang S Q, 2017. Cranial and dental material of Gomphotherium wimani (Gomphotheriidae, Proboscidea) from the Middle Miocene of the Linxia Basin, northwestern China. Vert PalAsiat, 55:331-346 |
| [166] | Zhang X Y, 1980. Fossil mammals from the Early Pleistocene of Yongren, Yunnan. Vert PalAsiat, 18:45-51 |
| [167] | Zhang X Y, 1982. The fossils of Pliocene elephantoidea from the basin of Lufeng, Yunnan. Vert PalAsiat, 20:359-368 |
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