欢迎访问《古脊椎动物学报》官方网站,今天是

鄂尔多斯盆地陕北肯氏兽组合带的肯氏兽型二齿兽类

展开
  • 1 中国科学院古脊椎动物与古人类研究所,中国科学院脊椎动物演化与人类起源重点实验室 北京 100044
    2 中国科学院大学地球与行星科学学院 北京 100049

收稿日期: 2022-01-27

  网络出版日期: 2022-06-01

基金资助

中国科学院国际合作局对外合作重点项目(132311KYSB20190010);中国科学院战略性先导科技专项(B类)(XDB26000000)

On kannemeyeriiform dicynodonts from the Shaanbeikannemeyeria Assemblage Zone of the Ordos Basin, China

Expand
  • 1 Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences Beijing 100044
    2 College of Earth and Planetary Sciences, University of Chinese Academy of Sciences Beijing 100049

Received date: 2022-01-27

  Online published: 2022-06-01

摘要

陕北肯氏兽(Shaanbeikannemeyeria)是二马营组下部常见的四足动物。这个属在分类学上还有一些问题没有解决,例如本属是否有效,包括几个种等。自1978年以来又陆续采集到一些材料。陕北肯氏兽首现于和尚沟组顶部。描述了发现的所有陕北肯氏兽化石,厘清其鉴定特征,了解个体差异,确认其系统位置。陕北肯氏兽只包含一个有效种:戏楼沟种(S. xilougouensis), 其独有衍征包括:枕部相对腭部强烈倾斜因而头骨腹面短,前颌骨有剑尖状的后背突,齿骨背侧内缘呈背凸的刀片状,隅骨的反折翼有一独立的后腹突,15枚背椎。这一属种在头骨形态上存在个体间差异,例如枕高相对枕宽比、吻尖形态(尖或钝)、颧弓眼眶部分形状以及齿突形态。有些差异可能与个体发育有关,例如齿突和长牙的发育程度、前颌骨在颧弓上向后延伸程度、前颌骨与额骨的距离、间颞部宽度及顶骨出露程度。基于头后骨骼可以判定二马营组下部还存在第二种肯氏兽类,可能是副肯氏兽(Parakannemeyeria)。

本文引用格式

刘俊 . 鄂尔多斯盆地陕北肯氏兽组合带的肯氏兽型二齿兽类[J]. 古脊椎动物学报, 2022 , 60(3) : 212 -248 . DOI: 10.19615/j.cnki.2096-9899.220601

Abstract

Shaanbeikannemeyeria is a common tetrapod from the lower part of the Ermaying Formation of the Ordos Basin, China. There are taxonomical questions surrounding this genus, such as the validity of the genus, and how many species are included within it. Several specimens have been collected since 1978. Shaanbeikannemeyeria first appeared from the top of the Heshanggou Formation. These specimens are described to clarify the diagnostic characters, the individual variations and the phylogenetic position of Shaanbeikannemeyeria. Only one species, S. xilougouensis, is recognized. It is characterized by the following autapomorphies: occiput strongly inclined relative to the palate such that the skull is much shorter basally than dorsally, sword tip-like premaxillary posterodorsal processes, tall and dorsally-convex cutting blade on the medial edge of the dorsal surface of the dentary, reflected lamina with a separated posteroventral process, and 15 dorsal vertebrae. This species shows variations on the cranial morphology, such as the occiput height relative to the width, the snout tip (sharp or obtuse), the shape of the orbital portion of the zygomatic arch, and the shape of caniniform process. Some variations could be ontogenetically related, such as the development of the caniniform process and tusk, the posterior extension of the maxilla on the zygomatic arch, the distance between the frontal and the premaxilla, the intertemporal bar width, and the exposing degree of the parietals. Based on postcranial bones, the second dicynodont genus (possibly Parakannemeyeria) is present in the lower Ermaying Formation.

参考文献

[1] Angielczyk K D, Hancox P J, Nabavizadeh A, 2017. A redescription of the Triassic kannemeyeriiform dicynodont Sangusaurus (Therapsida, Anomodontia), with an analysis of its feeding system. J Vert Paleont, 37(sup1): 189-227
[2] Angielczyk K D, Liu J, Yang W, 2021. A redescription of Kunpania scopulusa, a bidentalian dicynodont (Therapsida, Anomodontia) from the ?Guadalupian of northwestern China. J Vert Paleont, 41(1): e1922428
[3] Araújo D C, Gonzaga T D, 1980. Uma Nova especie de Jachaleria (Therapsida, Dicynodontia) do Triassico do Brasil. Actas Segundo Congr Argent Paleontol Bioestratigr Primer Congr Latino Am Paleontol, 2(1): 159-174
[4] Bandyopadhyay S, 1988. A kannemeyeriid dicynodont from the Middle Triassic Yerrapalli Formation. Philos Trans R Soc Lond B Biol Sci, 320: 185-233
[5] Bandyopadhyay S, 1989. The mammal-like reptile Rechnisaurus from the Triassic of India. Palaeontology, 32(2): 305-312
[6] Bonaparte J F, 1969. Dos nuevas “faunas” de reptiles triásicos de Argentina. In: Amos A J ed.ed. Gondwana Stratigraphy, IUGS Symposium, Buenos Aires, 1-15. Paris: United Nations Educational Scientific and Cultural Organization. 283-306
[7] Cheng Z W, 1980. 7. Vertebrate fossils. In: IGCAGS ed.ed. Mesozoic Stratigraphy and Palaeontology of the Shaanxi-Gansu-Ninxia Basin. Beijing: Geological Publishing House. 115-188
[8] Cox C B, 1959. On the anatomy of a new dicynodont genus with evidence of the position of the tympanum. Proc Zool Soc London, 132(3): 321-367
[9] Cox C B, 1968. The Chañares (Argentina) Triassic reptile fauna; IV, The dicynodont fauna. Breviora, 295: 1-27
[10] Cox C B, 1969. Two new dicynodonts from the Triassic Ntawere Formation, Zambia. Bull Br Mus Nat Hist Geol, 17: 257-294
[11] Cox C B, 1991. The Pangaea Dicynodont Rechnisaurus and the comparative biostratigraphy of Triassic dicynodont faunas. Palaeontology, 34: 767-784
[12] Cox C B, Li J L, 1983. A new genus of Triassic dicynodont from East Africa and its classification. Palaeontology, 26: 389-406
[13] Cruickshank A R I, 1967. A new Dicynodont genus from the Manda Formation of Tanzania (Tanganyika). J Zool, 153: 163-208
[14] Damiani R J, Vasconcelos C, Renaut A et al., 2007. Dolichuranus primaevus (Therapsida: Anomodontia) from the Middle Triassic of Namibia and its phylogenetic relationships. Palaeontology, 50: 1531-1546
[15] Danilov A I, 1971. A new dicynodont from the Middle Triassic of southern Cisuralia. Paleontol J, 5(2): 265-268
[16] Domnanovich N S, Marsicano C A, 2012. The Triassic dicynodont Vinceria (Therapsida, Anomodontia) from Argentina and a discussion on basal Kannemeyeriiformes. Geobios, 45(2): 173-186
[17] Dutuit J M, 1988. Ostéologie crânienne et ses enseignements, apports géologique et paléoécologique, de Moghreberia nmachouensis, Dicynodonte (Reptilia, Therapsida) du Trias supérieur marocain. Bull Mus Natn Hist Nat, 3: 227-285
[18] Goloboff P A, Catalano S, 2016. TNT, version 1.5, with a full implementation of phylogenetic morphometrics. Cladistics, 32: 221-238
[19] Govender R, 2005. Morphological and functional analysis of the postcranial anatomy of two dicynodont morphotypes from the Cynognathus assemblage zone of South Africa and their taxonomic implications. Ph. D thesis. Johannesburg: University of the Witwatersrand. 1-199
[20] Kalandadze N N, 1970. New Triassic kannemeyeriids from South Cisuralia. In: Flerov K K ed.ed. Materials on the Evolution of Terrestrial Vertebrates. Moscow: Nauka. 51-57
[21] Kammerer C F, 2018. The first skeletal evidence of a dicynodont from the lower Elliot Formation of South Africa. Palaeontol Afr, 52: 102-128
[22] Kammerer C F, Ordoñez M d l A, 2021. Dicynodonts (Therapsida: Anomodontia) of South America. J South Am Earth Sci, 108: 103171
[23] Kammerer C F, Fröbisch J, Angielczyk K D, 2013. On the validity and phylogenetic position of Eubrachiosaurus browni, a kannemeyeriiform dicynodont (Anomodontia) from Triassic North America. PLoS ONE, 8(5): e64203
[24] Kammerer C F, Viglietti P A, Hancox P J et al., 2019. A new kannemeyeriiform dicynodont (Ufudocyclops mukanelai, gen. et sp. nov.) from Subzone C of the Cynognathus Assemblage Zone, Triassic of South Africa, with implications for biostratigraphic correlation with other African Triassic Faunas. J Vert Paleont, 39(2): e1596921
[25] King G M, 1981. The postcranial skeleton of Robertia broomiana, an early dicynodont (Reptilia, Therapsida) from the South African Karoo. Ann S Afr Mus, 84(5): 203-231
[26] King G M, 1988. Anomodontia. Stuttgart: Gustav Fischer Verlag. 1-174
[27] Li J L, 1980. Kannemeyeria fossil from Inner Mongolia. Vert PalAsiat, 18(2): 94-99
[28] Li J L, Cheng Z W, 1995. A new Late Permian vertebrate fauna from Dashankou, Gansu, with comments on Permian and Triassic vertebrate assemblage zones of China. In: Sun A L, Wang Y Q eds. Sixth Symposium on Mesozoic Terrestrial Ecosystems and Biota, Short Papers. Beijing: China Ocean Press. 33-37
[29] Li J L, Liu J, 2015. Basal Synapsids. Palaeovertebrate Sinica, III(14): 1-105
[30] Liu J, 2015. New discoveries from the Sinokannemeyeria-Shansisuchus Assemblage Zone: 1. Kannemeyeriiformes from Shanxi, China. Vert PalAsiat, 53(1): 16-28
[31] Liu J, 2018. New progress on the correlation of Chinese terrestrial Permo-Triassic strata. Vert PalAsiat, 56(4): 327-342
[32] Liu J, 2021. The tetrapod fauna of the upper Permian Naobaogou Formation of China: 6. Turfanodon jiufengensis sp. nov. (Dicynodontia). Peer J, 9: e10854
[33] Liu J, Li J L, 2003. A new material of kannemeyerid from Xinjiang and the restudy of Parakannemeyeria brevirostris. Vert PalAsiat, 41(2): 147-156
[34] Liu J, Ramezani J, Li L et al., 2018. High-precision temporal calibration of Middle Triassic vertebrate biostratigraphy: U-Pb zircon constraints for the Sinokannemeyeria Fauna and Yonghesuchus. Vert PalAsiat, 56(1): 16-24
[35] Lucas S G, 2001. Chinese Fossil Vertebrates. New York: Columbia University Press. 1-375
[36] Maisch M W, 2001. Observations on Karoo and Gondwana vertebrates. Part 2: A new skull-reconstruction of Stahleckeria potens von Huene, 1935 (Dicynodontia, Middle Triassic) and a reconsideration of kannemeyeriiform phylogeny. Neues Jahrb Geol Palaontol Abh, 220(1): 127-152
[37] Morato L, 2006. Dinodontosaurus (Synapsida, Dicynodontia) reconstituições morfológicas e aspectos biomecânicos. Master thesis. Porto Alegre (RS): Universidade Federal do Rio Grande do Sul. 1-158
[38] Olson E C, 1944. Origin of mammals based upon cranial morphology of the therapsid suborders. Geol Soc Am Spec Pap, 55: 1-136
[39] Owen R, 1860. On the orders of fossil and recent Reptilia, and their distribution in time. Report of the Twenty-Ninth Meeting of the British Association for the Advancement of Science, 1859: 153-166
[40] Renaut A J, Hancox P J, 2001. Cranial description and taxonomic re-evaluation of Kannemeyeria argentinensis (Therapsida: Dicynodontia). Palaeontol Afr, 37: 81-91
[41] Sullivan C, Reisz R R, 2005. Cranial anatomy and taxonomy of the Late Permian dicynodont Diictodon. Ann Carnegie Mus, 74(1): 45-75
[42] Sun A L, 1960. On a new genus of kannemeyerids from Ningwu, Shanxi. Vert PalAsiat, 4(2): 67-80
[43] Sun A L, 1963. The Chinese kannemeyerids. Palaeont Sin, New Ser C, 17: 1-109
[44] Sun A L, 1978. Discovery of Parakannemeyeria from Xinjiang. Mem Inst Vert Paleont Paleoanthrop, Acad Sin, 13: 47-54
[45] Surkov M V, 2003. A new anomodont (Therapsida) from the Middle Triassic of the southern Fore-Urals. Paleontol J, 37(4): 425-431
[46] Vega-Dias C, Schultz C L, 2004. Postcranial material of Jachaleria candelariensis Araújo and Gonzaga 1980 (Therapsida, Dicynodontia), Upper Triassic of Rio Grande do Sul, Brazil. PaleoBios, 24(1): 7-31
[47] Vega-Dias C, Maisch M W, Schwanke C, 2005. The taxonomic status of Stahleckeria impotens (Therapsida, Dicynodontia): redescription and discussion of its phylogenetic position. Rev Bras Paleontol, 8(3): 221-228
[48] von Huene F F, 1935. Die fossilen Reptilien des südamerikanischen Gondwanalandes. Ergebnisse der Sauriergrabungen in Südbrasilien, 1928-1929. Ordnung Anomodontia. Munich: C. H. Beck’sche Verlagsbuchhandlung. 1-92
[49] Williston S W, 1904. Notice of some new reptiles from the Upper Trias of Wyoming. J Geol, 12(8): 688-697
[50] Young C C, 1937. On the Triassic dicynodonts from Shansi. Bull Geol Soc China, 17: 393-411
文章导航

/