Vertebrata Palasiatica ›› 2021, Vol. 59 ›› Issue (4): 273-294.DOI: 10.19615/j.cnki.2096-9899.210715
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DONG Wei1,2(), BAI Wei-Peng1,2,3, ZHANG Li-Min1,2,3
Received:
2021-03-26
Online:
2021-10-20
Published:
2021-10-18
通讯作者:
* dongwei@ivpp.ac.cn基金资助:
CLC Number:
DONG Wei, BAI Wei-Peng, ZHANG Li-Min. The first description of Rhinocerotidae (Perissodactyla, Mammalia) from Xinyaozi Ravine in Shanxi, North China. Vertebrata Palasiatica, 2021, 59(4): 273-294.
董为, 白炜鹏, 张立民. 2021, 59(4): 273-294, 山西天镇辛窑子早更新世犀科化石新材料. 古脊椎动物学报.
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URL: https://www.vertpala.ac.cn/EN/10.19615/j.cnki.2096-9899.210715
Taxon | Elasmotherium peii | E. cf. E. peii | E. caucasicum | E. sibiricum | ||||||
---|---|---|---|---|---|---|---|---|---|---|
Location | Xinyaozi | Shanshenmiaozhui1) | Xihoudu2) | Stavropol Krai3) | Salcia4) | Taman5) | Stavropol Krai3) | |||
m2 L | 74.44 | 75 | 75.5 | 70-77 | 70-77 | 70-95 | 64.2-84 | |||
m2 W | 40.58 | 40 | 43 | 41.5-60 | 34-36 | 39-50 | 34-43.5 | |||
m2 H | 29.12(92) | 104 | 44-77 | |||||||
m3 L | 82.36 | 72-84 | 60-98 | |||||||
m3 W | 37.78 | 33 | 30-44 | |||||||
m3 H | 31.28(140) | 129-130 |
Taxon | Elasmotherium peii | E. cf. E. peii | E. caucasicum | E. sibiricum | ||||||
---|---|---|---|---|---|---|---|---|---|---|
Location | Xinyaozi | Shanshenmiaozhui1) | Xihoudu2) | Stavropol Krai3) | Salcia4) | Taman5) | Stavropol Krai3) | |||
m2 L | 74.44 | 75 | 75.5 | 70-77 | 70-77 | 70-95 | 64.2-84 | |||
m2 W | 40.58 | 40 | 43 | 41.5-60 | 34-36 | 39-50 | 34-43.5 | |||
m2 H | 29.12(92) | 104 | 44-77 | |||||||
m3 L | 82.36 | 72-84 | 60-98 | |||||||
m3 W | 37.78 | 33 | 30-44 | |||||||
m3 H | 31.28(140) | 129-130 |
Fig. 2 A right juvenile maxillary fragment (IVPP V 27429.1) of Stephanorhinus cf. S. kirchbergensis from Xinyaozi Ravine in lingual (A), occlusal (B) and buccal (C) views
V 27429.1 | Xiashagou1) | Rhino Cave2) | Huludong3) | Zhoukoudian4) Loc. 1 | Zhoukoudian5) Loc. 13 | West Europe6) | |
---|---|---|---|---|---|---|---|
DP1 L | 25.38 | 26 | 31.8-32.9 | 28-29 | 27.5 | 35 | 26-32 |
DP1 W | 20.06 | 24 | 26-27.9 | 28 | 24-25 | 32 | 31-34 |
DP1 H | 24.34 | 26 | |||||
DP2 L | 35.08 | 37 | 44-47.3 | 38-39 | 36.5 | 41 | 34-39.5 |
DP2 W | 32.76 | 40 | 42.6-47.9 | 40-42 | 39.5 | 40 | 31-43 |
DP2 H | 28.48 | 25 | |||||
DP3 L | 41.08 | 50.6-55.8 | 47-48 | 43.5 | 51 | 41.5-50.5 | |
DP3 W | 39.12 | 48.2-52 | 50-52 | 49 | 53 | 42-54.5 | |
DP3 H | 35.18 | 36.5 | |||||
DP4 L | 49.52 | 52 | 56.6-60 | 48-52 | 47 | 53 | 48-57 |
DP4 W | 34.24 | 51 | 52.6-54.5 | 54 | 48.5 | 56 | 51-57.5 |
DP4 H | 42.56 | 42 | |||||
L DP1-4 | 145.32 | 172.8-183 | 175-177 | 147 | 150 |
V 27429.1 | Xiashagou1) | Rhino Cave2) | Huludong3) | Zhoukoudian4) Loc. 1 | Zhoukoudian5) Loc. 13 | West Europe6) | |
---|---|---|---|---|---|---|---|
DP1 L | 25.38 | 26 | 31.8-32.9 | 28-29 | 27.5 | 35 | 26-32 |
DP1 W | 20.06 | 24 | 26-27.9 | 28 | 24-25 | 32 | 31-34 |
DP1 H | 24.34 | 26 | |||||
DP2 L | 35.08 | 37 | 44-47.3 | 38-39 | 36.5 | 41 | 34-39.5 |
DP2 W | 32.76 | 40 | 42.6-47.9 | 40-42 | 39.5 | 40 | 31-43 |
DP2 H | 28.48 | 25 | |||||
DP3 L | 41.08 | 50.6-55.8 | 47-48 | 43.5 | 51 | 41.5-50.5 | |
DP3 W | 39.12 | 48.2-52 | 50-52 | 49 | 53 | 42-54.5 | |
DP3 H | 35.18 | 36.5 | |||||
DP4 L | 49.52 | 52 | 56.6-60 | 48-52 | 47 | 53 | 48-57 |
DP4 W | 34.24 | 51 | 52.6-54.5 | 54 | 48.5 | 56 | 51-57.5 |
DP4 H | 42.56 | 42 | |||||
L DP1-4 | 145.32 | 172.8-183 | 175-177 | 147 | 150 |
Fig. 3 A broken mandible (IVPP V 27429.3) of Stephanorhinus cf. S. kirchbergensis from Xinyaozi Ravine in buccal (A), occlusal (B) and lingual (C) views
Fig. 4 Mandibular fragments (IVPP V 27429.4-5) of Stephanorhinus cf. S. kirchbergensis from Xinyaozi Ravine A-C. V 27429.4; D-F. V 27429.5; A, D. lingual view; B, E. occlusal view; C. buccal view; F. sagittal slice of CT image
V 27429.3 | V 27429.4 | V 27429.5 | Rhino Cave1) | Loc. 13 Zhoukoudian2) | West Europe3) | |
---|---|---|---|---|---|---|
dp1 L | 20.56 | 23-23.4 | 21 | 17.5-23 | ||
dp1 W | 11.38 | 11.6-13 | 13 | 14 | ||
dp1 H | 19.82 | 18.5 | ||||
dp2 L | 32.42 | 32.32 | 30.5-38.8 | 35 | 28-38 | |
dp2 W | 18.44 | 18.94 | 16.7-20 | 21 | 16.5-21.5 | |
dp2 H | 25.02 | 25.12 | ||||
dp3 L | 44.12 | 46.98 | 45.78 | 49.2 | 44 | 38-44.5 |
dp3 W | 23.22 | 24.98 | 25.44 | 25.1 | 26 | 21-28.5 |
dp3 H | 28.84 | 30.26 | 32.82 | 28.5 | ||
dp4 L | 44.82 | 45.18 | 47.68 | 49.5-52 | 47 | 42-51 |
dp4 W | 23.22 | 27.46 | 21.54 | 27.1-27.4 | 29 | 21-31 |
dp4 H | 32.12 | 34.14 | 23.14 | 30-35.5 | ||
L dp1-4 | 141.68 | 182-201 |
V 27429.3 | V 27429.4 | V 27429.5 | Rhino Cave1) | Loc. 13 Zhoukoudian2) | West Europe3) | |
---|---|---|---|---|---|---|
dp1 L | 20.56 | 23-23.4 | 21 | 17.5-23 | ||
dp1 W | 11.38 | 11.6-13 | 13 | 14 | ||
dp1 H | 19.82 | 18.5 | ||||
dp2 L | 32.42 | 32.32 | 30.5-38.8 | 35 | 28-38 | |
dp2 W | 18.44 | 18.94 | 16.7-20 | 21 | 16.5-21.5 | |
dp2 H | 25.02 | 25.12 | ||||
dp3 L | 44.12 | 46.98 | 45.78 | 49.2 | 44 | 38-44.5 |
dp3 W | 23.22 | 24.98 | 25.44 | 25.1 | 26 | 21-28.5 |
dp3 H | 28.84 | 30.26 | 32.82 | 28.5 | ||
dp4 L | 44.82 | 45.18 | 47.68 | 49.5-52 | 47 | 42-51 |
dp4 W | 23.22 | 27.46 | 21.54 | 27.1-27.4 | 29 | 21-31 |
dp4 H | 32.12 | 34.14 | 23.14 | 30-35.5 | ||
L dp1-4 | 141.68 | 182-201 |
Fig. 5 Broken mandible (IVPP V 27429.2) of Stephanorhinus cf. S. kirchbergensis from Taijiaping in Xinyaozi Ravine in lingual (A), occlusal (B) and buccal (C) views
IVPP V 27429.2 | S. kirchbergensis | |||||||
---|---|---|---|---|---|---|---|---|
left | right | Nihewan1) | Rhino Cave2) | Loc. 1 Zhoukoudian3) | Anping4) | West Europe5) | ||
p2 L | 28.66 | 26.88 | 26.7-29.2 | 30.5-33.5 | 26-32 | 29.1-32.7 | 30.5-37 | |
p2 W | 17.06 | 17.04 | 17.2-18 | 18.7-20.5 | 17-23 | 18.5-21.8 | 18.5-23 | |
p2 H | 21.72 | 19.18 | 20-36 | 28-40 | ||||
p3 L | 32.56 | 31.14 | 26.3 | 34.5-36.8 | 33-38 | 32.6-41.1 | 40-46.5 | |
p3 W | 22.84 | 22.48 | 24 | 25.9-28.6 | 21-29 | 26.5-28.4 | 23.5-30 | |
p3 H | 20.58 | 18.52 | 17-32 | 36 | ||||
p4 L | 35.88 | 35.12 | 38-38.4 | 42-43.4 | 40-49 | 37.9-42.8 | 39-49 | |
p4 W | 25.02 | 25.36 | 28-28.6 | 30-32.7 | 30-36 | 29.6-33.7 | 25.5-36 | |
p4 H | 22.68 | 21.92 | 25-34 | 28.5-53 | ||||
m1 L | 39.34 | 37.52 | 43-43.3 | 41.8-47.9 | 40-45 | 44.1-51.1 | 46.5-56 | |
m1 W | 30.28 | 29.18 | 31.5-31.9 | 33.1-37 | 29-33 | 30.8-37.1 | 27-37 | |
m1 H | 15.98 | 15.82 | 17-31 | 40-58 | ||||
m2 L | 46.86 | 42.42 | 48-48.7 | 51.8-56.5 | 38-53 | 49.5-54.7 | 51-63 | |
m2 W | 31.92 | 32.08 | 30.9-31.8 | 34.6-36.3 | 29-34 | 32.1-36.3 | 30-40 | |
m2 H | 23.52 | 22.08 | 22-49 | 42-54 | ||||
m3 L | 47.02 | 48.76 | 46.3-46.8 | 52.1-60.4 | 47-57 | 52.1-55.8 | 52-68 | |
m3 W | 28.24 | 29.08 | 29.4-29.5 | 32.6-35.7 | 30-38 | 31.7-32.8 | 28-40 | |
m3 H | 26.08 | 26.02 | 39-47 | 40-64 | ||||
L p2-4 | 89.62 | 88.42 | ||||||
L m1-3 | 130.42 | 129.08 | 130-137 | |||||
L p2-m3 | 219.96 | 214.38 | 248.1 |
IVPP V 27429.2 | S. kirchbergensis | |||||||
---|---|---|---|---|---|---|---|---|
left | right | Nihewan1) | Rhino Cave2) | Loc. 1 Zhoukoudian3) | Anping4) | West Europe5) | ||
p2 L | 28.66 | 26.88 | 26.7-29.2 | 30.5-33.5 | 26-32 | 29.1-32.7 | 30.5-37 | |
p2 W | 17.06 | 17.04 | 17.2-18 | 18.7-20.5 | 17-23 | 18.5-21.8 | 18.5-23 | |
p2 H | 21.72 | 19.18 | 20-36 | 28-40 | ||||
p3 L | 32.56 | 31.14 | 26.3 | 34.5-36.8 | 33-38 | 32.6-41.1 | 40-46.5 | |
p3 W | 22.84 | 22.48 | 24 | 25.9-28.6 | 21-29 | 26.5-28.4 | 23.5-30 | |
p3 H | 20.58 | 18.52 | 17-32 | 36 | ||||
p4 L | 35.88 | 35.12 | 38-38.4 | 42-43.4 | 40-49 | 37.9-42.8 | 39-49 | |
p4 W | 25.02 | 25.36 | 28-28.6 | 30-32.7 | 30-36 | 29.6-33.7 | 25.5-36 | |
p4 H | 22.68 | 21.92 | 25-34 | 28.5-53 | ||||
m1 L | 39.34 | 37.52 | 43-43.3 | 41.8-47.9 | 40-45 | 44.1-51.1 | 46.5-56 | |
m1 W | 30.28 | 29.18 | 31.5-31.9 | 33.1-37 | 29-33 | 30.8-37.1 | 27-37 | |
m1 H | 15.98 | 15.82 | 17-31 | 40-58 | ||||
m2 L | 46.86 | 42.42 | 48-48.7 | 51.8-56.5 | 38-53 | 49.5-54.7 | 51-63 | |
m2 W | 31.92 | 32.08 | 30.9-31.8 | 34.6-36.3 | 29-34 | 32.1-36.3 | 30-40 | |
m2 H | 23.52 | 22.08 | 22-49 | 42-54 | ||||
m3 L | 47.02 | 48.76 | 46.3-46.8 | 52.1-60.4 | 47-57 | 52.1-55.8 | 52-68 | |
m3 W | 28.24 | 29.08 | 29.4-29.5 | 32.6-35.7 | 30-38 | 31.7-32.8 | 28-40 | |
m3 H | 26.08 | 26.02 | 39-47 | 40-64 | ||||
L p2-4 | 89.62 | 88.42 | ||||||
L m1-3 | 130.42 | 129.08 | 130-137 | |||||
L p2-m3 | 219.96 | 214.38 | 248.1 |
Fig. 6 Length range comparison of lower cheek teeth from selected localities Abbreviations and data sources: ZKD Loc9, Coelodonta antiquitatis from Locality 9 at Zhoukoudian (Teilhard de Chardin, 1936); Co W Eu, C. antiquitatis from West Europe (Guérin, 1980); LD, C. nihowanensis from Longdan (Qiu et al., 2004); St W Eu, Stephanorhinus kirchbergensis from West Europe (Guérin, 1980); ZKD Loc1, S. k. from Locality 1 of Zhoukoudian (Chow, 1979); RhinoCave, S. k. from Shennongjia (Tong and Wu, 2010); DHG-SG, S. k. from Daheigou and Shigou in Nihewan Basin (Tong et al., 2014); XYZ, Xinyaozi Ravine
V 27430.1 | C. nihowanensis | C. antiquitatis | C. tologoijensis | |||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|
left | right | Xiashagou1) | Danangou2) | SSMZ3) | Longdan4) | Dingcun5) | W. Europe6) | W. Transbaikalia7) | ||||
DP1 L | 29.48 | 29.04 | 32 | 27 | 26-27 | 27.3 | 26 | 21-21.5 | 22-27 | |||
DP1 W | 22.04 | 21.68 | 23 | 20 | 18.2-20 | 21.8 | 19 | 17-18 | 17-20 | |||
DP1 H | 21.34 | 22.64 | ||||||||||
DP2 L | 34.78 | 34.32 | 35 | 34.8 | 31.5-32.5 | 34.5 | 27.5 | 28-33 | 28-32 | |||
DP2 W | 35.08 | 33.76 | 34 | 32 | 29.2-30.3 | 35 | 32 | 26-32.5 | 28-30 | |||
DP2 H | 26.58 | 27.56 | 23-30 | |||||||||
DP3 L | 45.82 | 46.46 | 47 | 45 | 42.6-44.7 | 41.7 | 40-48 | 31-45.5 | 43-49 | |||
DP3 W | 41.02 | 40.18 | 40 | 39 | 37.6-38.3 | 40.8 | 37-45 | 31.5-42 | 35-41 | |||
DP3 H | 34.22 | 35.28 | 25-33.5 | |||||||||
DP4 L | 49.54 | 54 | 46.9 | 50.2 | 55-55.5 | 44-57 | ||||||
DP4 W | 43.46 | 40 | 41.4 | 46 | 40-43 | 39-55 | ||||||
DP4 H | 35.28 | 33.5-46 | ||||||||||
L DP1-4 | 145.62 | 152 | 136 | 150 | 128-141 |
V 27430.1 | C. nihowanensis | C. antiquitatis | C. tologoijensis | |||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|
left | right | Xiashagou1) | Danangou2) | SSMZ3) | Longdan4) | Dingcun5) | W. Europe6) | W. Transbaikalia7) | ||||
DP1 L | 29.48 | 29.04 | 32 | 27 | 26-27 | 27.3 | 26 | 21-21.5 | 22-27 | |||
DP1 W | 22.04 | 21.68 | 23 | 20 | 18.2-20 | 21.8 | 19 | 17-18 | 17-20 | |||
DP1 H | 21.34 | 22.64 | ||||||||||
DP2 L | 34.78 | 34.32 | 35 | 34.8 | 31.5-32.5 | 34.5 | 27.5 | 28-33 | 28-32 | |||
DP2 W | 35.08 | 33.76 | 34 | 32 | 29.2-30.3 | 35 | 32 | 26-32.5 | 28-30 | |||
DP2 H | 26.58 | 27.56 | 23-30 | |||||||||
DP3 L | 45.82 | 46.46 | 47 | 45 | 42.6-44.7 | 41.7 | 40-48 | 31-45.5 | 43-49 | |||
DP3 W | 41.02 | 40.18 | 40 | 39 | 37.6-38.3 | 40.8 | 37-45 | 31.5-42 | 35-41 | |||
DP3 H | 34.22 | 35.28 | 25-33.5 | |||||||||
DP4 L | 49.54 | 54 | 46.9 | 50.2 | 55-55.5 | 44-57 | ||||||
DP4 W | 43.46 | 40 | 41.4 | 46 | 40-43 | 39-55 | ||||||
DP4 H | 35.28 | 33.5-46 | ||||||||||
L DP1-4 | 145.62 | 152 | 136 | 150 | 128-141 |
C. nihowanensis | C. antiquitatis | C. tologoijensis | ||||||||
---|---|---|---|---|---|---|---|---|---|---|
V 27430.2 | V 27430.3 | Xiashagou1) | Danangou2) | Shanshenmiaozui3) | Dingcun4) | W. Europe5) | W. Transbaikalia6) | |||
dp3 L | 40.08 | 38 | 41 | 38.6-40 | 37-38.2 | 34.5-41.5 | 36-38 | |||
dp3 W | 19.98 | 18 | 21 | 21-23 | 19.5-21.3 | 17-24 | 16-18 | |||
dp3 H | 32.76 | 24.5-36 | ||||||||
dp4 L | 45.12 | 46 | 44 | 41.5-45 | 45-23 | 37-51 | 39-43 | |||
dp4 W | 24.02 | 22 | 21 | 21.2-25 | 19-25.5 | 19-21 | ||||
dp4 H | 32.66 | 27-46 |
C. nihowanensis | C. antiquitatis | C. tologoijensis | ||||||||
---|---|---|---|---|---|---|---|---|---|---|
V 27430.2 | V 27430.3 | Xiashagou1) | Danangou2) | Shanshenmiaozui3) | Dingcun4) | W. Europe5) | W. Transbaikalia6) | |||
dp3 L | 40.08 | 38 | 41 | 38.6-40 | 37-38.2 | 34.5-41.5 | 36-38 | |||
dp3 W | 19.98 | 18 | 21 | 21-23 | 19.5-21.3 | 17-24 | 16-18 | |||
dp3 H | 32.76 | 24.5-36 | ||||||||
dp4 L | 45.12 | 46 | 44 | 41.5-45 | 45-23 | 37-51 | 39-43 | |||
dp4 W | 24.02 | 22 | 21 | 21.2-25 | 19-25.5 | 19-21 | ||||
dp4 H | 32.66 | 27-46 |
Fig. 8 Deciduous cheek teeth of Coelodonta nihowanensis from Xinyaozi Ravine A-C. right dp3 (IVPP V 27430.2); D-F. right dp4 (V 27430.3) A, D. buccal view; B, E. occlusal view; C, F. lingual view
Fig. 9 Ratio diagram of dimensions of upper deciduous cheek teeth from related localities, plotted against those of IVPP V 27430.1 (reference 0) Note: Data source as in Tables 2, 5. L. length; W. width; ZKD. Zhoukoudian; solid line, Coelodonta nihowanensis ; dash line, C. antiquitatis ; dotted line, Stephanorhinus kirchbergensis
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