The tetrapod fauna of the upper Permian Naobaogou Formation of China— 4. the diversity of dicynodonts

doi:10.19615/j.cnki.1000-3118.190522

Abstract

Abstract:

The Permian dicynodont records were previously rare in North China, but many dicynodont specimens have been collected from the Naobaogou Formation in Daqingshan area, Nei Mongol since 2009. Among these specimens, seven morphotypes have been identified, and they may represent seven different species: two of them are closely related to Daqingshanodon limbus, three or four are closely related to Jimusaria sinkianensis, and one may be closely related to Turfanodon. This study shows the dicynodonts also have a high diversity at the species level in North China.

Key words: Daqingshan area,Nei Mongol, Permian, Naobaogou Formation, Daqingshanodon, Jimusaria, Turfanodon

摘要：

华北二叠纪的二齿兽类记录很少。从2009年开始在内蒙古大青山地区脑包沟组中采集到大量的二齿兽类化石,从中可以鉴别出7个形态类型,它们可能代表了7个物种：其中2个与边缘大青山兽(Daqingshanodon limbus)密切相关,3或4个与新疆吉木萨尔兽(Jimusaria sinkianensis)关系密切,1个可能与吐鲁番兽(Turfanodon)关系密切。研究表明华北的二齿兽类在种级也具有很高的多样性。

关键词: 内蒙古大青山, 二叠纪, 脑包沟组, 大青山兽, 吉木萨尔兽, 吐鲁番兽

CLC Number:

Q915.861

LIU Jun. The tetrapod fauna of the upper Permian Naobaogou Formation of China— 4. the diversity of dicynodonts. Vertebrata Palasiatica, 2019, 57(3): 173-180.

刘俊. 2019, 57(3): 173-180, 二叠纪脑包沟组的四足动物群——4. 二齿兽类的多样性. 古脊椎动物学报.

Figures/Tables 4

References 23

[1]	Amalitzky V, 1922. Diagnoses of the new forms of vertebrates and plants from the Upper Permian on North Dvina. Bull Acad Sci Russ, VI Ser, 16:329-340
[2]	Angielczyk K D, Kammerer C F, 2017. The cranial morphology, phylogenetic position and biogeography of the upper Permian dicynodont Compsodon helmoedi van Hoepen (Therapsida, Anomodontia). Pap Palaeontol, 3(4):513-545 DOI URL
[3]	Bernardi M, Petti F M, Kustatscher E et al., 2017. Late Permian (Lopingian) terrestrial ecosystems: a global comparison with new data from the low-latitude Bletterbach Biota. Earth-Sci Rev, 175(Suppl C):18-43 DOI URL
[4]	Kammerer C F, Angielczyk K D, Fröbisch J, 2011. A comprehensive taxonomic revision of Dicynodon (Therapsida, Anomodontia) and its implications for dicynodont phylogeny, biogeography, and biostratigraphy. J Vert Paleont, 31(suppl 1):1-158
[5]	Kammerer C F, Angielczyk K D, Fröbisch J, 2015. Redescription of the geikiid Pelanomodon (Therapsida, Dicynodontia), with a reconsideration of ‘Propelanomodon’. J Vert Paleont, 36(1):e1030408 DOI URL
[6]	King G M, 1988. Anomodontia, Encyclopedia of Paleoherpetology. Stuttgart: Gustav Fischer Verlag. 174
[7]	Kurkin A A, 1999. A new dicynodont from the Malaya Severnaya Dvina River excavations. Paleontol J, 33:297-301
[8]	Kurkin A A, 2000. New dicynodonts from the Upper Permian of the Vyatka Basin. Paleontol J, 34:203-210
[9]	Kurkin A A, 2001. New Late Permian dicynodonts from the Vyazniki assemblage of terrestrial tetrapods of Eastern Europe. Paleontol J, 35:53-59
[10]	Kurkin A A, 2006. A new dicynodont (Anomodontia, Eotherapsida) from the Upper Permian of Tatarstan. Paleontol J, 40:434-437 DOI URL
[11]	Kurkin A A, 2012. Dicynodontids of Eastern Europe. Paleontol J, 46:187-198 DOI URL
[12]	Li J L, Liu J, 2015. Palaeovertebrata Sinica, Vol 3, Ser 14 Basal Synapsids. Beijing: Science Press. 1-105
[13]	Li P X, Cheng Z W, Li J L, 2000. A new species of Dicynodon from Upper Permian of Sunan, Gansu, with remarks on related strata. Vert PalAsiat, 38(2):147-157
[14]	Liu J, Abdala F, 2017. The tetrapod fauna of the upper Permian Naobaogou Formation of China: 1. Shiguaignathus wangi gen. et sp. nov., the first akidnognathid therocephalian from China. PeerJ, 5:e4150 DOI URL
[15]	Liu J, Abdala F, 2019. The tetrapod fauna of the upper Permian Naobaogou Formation of China: 3. Jiufengia jiai gen. et sp. nov., a large akidnognathid therocephalian. PeerJ, 7:e6463 DOI URL
[16]	Liu J, Bever Gabriel S, 2018. The tetrapod fauna of the upper Permian Naobaogou Formation of China: a new species of Elginia (Pareiasauria). Pap Palaeontol, 4(2):197-209 DOI URL
[17]	Sun A L, 1963. The Chinese kannemeyerids. Palaeontol Sin, New Ser C, 17:1-109
[18]	Sun A L, 1973a. Permo-Triassic dicynodonts from Turfan, Sinkiang. Mem Inst Vert Paleont Paleoanthrop, Acad Sin, 10:53-68
[19]	Sun A L, 1973b. A new species of Dicynodon from Sinkiang. Vert PalAsiat, 11(1):52-58
[20]	Sun A L, 1978. Two new genera of Dicynodontidae. Mem Inst Vert Paleont Paleoanthrop, Acad Sin, 13:19-25
[21]	Sushkin P P, 1926. Notes on the Pre-Jurassic Tetrapoda from Russia. I. Dicynodon amalitzkii, n. sp. Palaeontol Hung, 1:323-327
[22]	Yuan P L, Young C C, 1934. On the discovery of a new dicynodon in Sinkiang. Bull Geol Surv China, 13:563-573
[23]	Zhu Y L, 1989. The discovery of dicynodonts in Daqingshan Montain, Nei Mongol (Inner Mongolia). Vert PalAsiat, 27(1):9-27

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IVPP V 23878	1	1	0	0	?	1	2	0	1	2	1
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